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ThefourthseasonofAllCreaturesGreatandSmallwillpickupinspringtimeof1940withchangeonthehorizonforeveryoneinSkeldaleHouse.JamesandHelenwonderwhentherighttimemightbetostartafamily,notknowingwhetherornotJameswillbecalleduptoserveintheRAF.Tristan’sabsenceisfeltbyall,butnoonemorethanSiegfriedwhoattemptstoholdthegrowinghousehold,andhimself,togetherashebravesthisnewworld.MrsHallandHelen’sfriendshipblossomsastheylooktothefutureandnewtraineeveterinarystudentRichardCarmodyarrives,causingcomplicationsinthehouse.